Dave (a.k.a Dr. Octavius Brine) is the main antagonist of Penguins of Madagascar: The Movie.
Dave was originally one of the star attractions of the Central Park Zoo, until Skipper, Kowalski, Rico and Private arrived and stole his fame with their cuteness, causing him to be roughly removed from his home. Afterwards, Dave was shipped from zoo to zoo only to have his fame stolen by other penguins. Because of this, he becomes embittered and seeks revenge on the penguin species, becoming a mad scientist in the process.
He is quite jealous of the penguins because he is always being overthrown by penguins. He has a type of revenge spirit for when he says I got to do some shopping, FOR REVENGE! he seems to have a type of villain spirit in him. He is also shown to be very poor in computers, for example when he called North Wind Headquarters and had problems turning on his mike, and putting his camera on and hanging up. Like Dr. Blowhole he has a hatred of the penguins but unlike Blowhole Dave doesn't seem to have any hatred for the humans.
Dave like all octopuses has the ability to disguise himself by changing his skin color and molding his bone-less body into different shapes. He was able to squeeze himself into a vending machine in order to capture Skipper, Kowalski, Rico, and Private. He usually disguises himself as a human geneticist that goes by the name Dr. Octavius Brine. Unlike his octopus henchmen, Dave conveniently has a freakish human-like face and mouth which lets him easily pass as a human after soing some minor disguise alterations. Similarity to the tv-show-exclusive villain Dr. Blowhole ,he is extremely intelligent. He was also able to hack into North Wind's computer systems.
- Medusa Serum- Serum that mutates anything into horrible monsters. Can be neutralized by something cute.
- Dave's Ray- Fires the medusa serum to mutate penguins. Later it's captured and detained by the penguins in order to revert Private void of his horns.
- Speech Translator microphone-Has a microphone that allows him to speak to humans so can continue pretending he's a human(since animals can't talk to humans in the Madagascar-verse)