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The Fossas (alternatively spelled Foosa) are the central antagonists of the Madagascar franchise. They are the main antagonists of Madagascar, and the tertiary antagonists of All Hail King Julien/All Hail King Julien: Exiled.

They appear as medium-sized feline-like predators that rarely speak, and instead they growl and yelp. They are described by Kowalski as "wild beasts that bite with flesh-shredding, bone-crushing jaws of doom.". They are dangerous, ruthless, evil, mean, wild, murderous, power-hungry, selfish, greedy and feral. One of the reasons why they are this way is because they want food.

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Throughout Madagascar, they are shown to terrorize the lemurs and attempt to eat them. According to Julien, "they're always annoying us by trespassing, interrupting our parties, and ripping our limbs off." When they do speak, they put "fossa" in front of every word: ex. "Fossa hungry, Fossa eat." They are terrified of Alex due to him being a lion and being much more physically powerful and intimidating. They begin to scare Marty, Gloria, and Melman next. The penguins stop them with a halt gun and used Private as an example, but Alex roars so loud that Fossas stop and copies the fossas' words by saying "Alex hungry, Alex eat". In the end, thanks to a setup performed by his friends and the penguins, Alex scares them away making sure that the fossa never bothered the lemurs ever again.

In The Penguins of Madagascar episode "The Terror of Madagascar," a baby fossa appears. It is innocent, but Julien thinks that he is trying to eat him. The fossa is revealed to think Julien as his father and saved him from Savio.

The Fossa appears as the tertiary antagonist in All Hail King Julien. In the past, King Julien’s uncle, who was the twelfth king of the lemurs, kept the kingdom silent in order to keep the fossa away until he was foretold that he would be eaten.


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